Abstract
1 Department of Environmental Ecology, Faculty of Environmental Science, Hanoi University of Science, VNU, 334 Nguyen Trai Road, Thanh Xuan District, Hanoi, Vietnam 2 Centre for Natural Resources and Environmental Studies, VNU, 19 Le Thanh Tong Street, Hanoi, Vietnam 3 Department of Genetics, Faculty of Biology, Hanoi University of Science, VNU, 334 Nguyen Trai Road, Thanh Xuan District, Hanoi, Vietnam 4 Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam 5 Fauna and Flora International, 340 Nghi Tam Road, Tay Ho District, Hanoi, Vietnam 6 Phong Nha-Ke Bang Conservation National Park, Quang Binh Province, Vietnam
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