Abstract
Phylogenetic trends of the states of 24 adult and 14 larval characters of the major groups of Scarabaeoidea are discussed on the basis of outgroup and ingroup comparison. Evolutionary trends within the major groups are also discussed. The following (i) adult and (ii) larval characters are discussed: (i) antennal segments, antennal sensilla, ocular canthus, ommatidium structure, epipharynx, mandibles, maxillae, labium, tentorium, trochantin, coxal cavities, mesothoracic spiracles, wing venation, metendosternite, empodium, abdominal sternites, abdominal spiracles, male genitalia, female genitalia, ovarioles, karyotype, central nervous system, spermatozoan number and malpighian tubules; (ii) antennae, frontoclypeal suture, lateral ocelli (stemmata), labial palpi, maxillae, maxillary palpi, maxillary/mandibular stridulatory areas, labrum, epipharyngeal tormae, legs, stridulatory apparatus, spiracles, abdominal apex and central nervous system.
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