Abstract
Institute of Ecology and Resource Management, Division of Biological Sciences, University of Edinburgh, Darwin Building, Kings Buildings, Mayfield Road, Edinburgh EH9 3JU, tDepartment of Forest Resources Management, University of Ibadan, Ibadan, Nigeria, tDepartment of Biology, University of Botswana, Private Bag 0022, Gaborone, Botswana and ?Forestry Research Institute of Nigeria, Ibadan, Nigeria
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