Abstract
In Nothofagus, the staminate inflorescences occur in the proximal leaf axils of proleptic seasonal growth units. The subgenera Brassospora, Fuscospora, and Nothofagus have either dichasia of three flowers or a single flower; subgenus Lophozonia has only solitary flowers. Flowers of Lophozonia differ significantly from those of the other subgenera. They have substantially more perianth lobes and stamens, stamen initiation follows a pseudocentrifugal pattern, and the floral meristem is distinctly oval in shape. The solitary “flower” of Lophozonia is interpreted as a complex structure: a “pseudanthium” formed by the failure to ramify of a three‐flowered dichasium. Pseudanthium development is a potential synapomorphy supporting the monophyly of subgenus Lophozonia (Turcz.) Krasser. Floral and inflorescence morphology in Nothofagus is compared with other families in the “higher” Hamamelididae.
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