Abstract

Abstract The genus Gnosonesima (Turbellaria, Lecithoepitheliata) is revised on the basis of all four known species, G. antarctica Reisinger, 1926, G. borealis (Westblad, 1952), G. brattstroemi sp. n., and G. reisingeri sp. n. It is the only genus in the family Gnosonesimidae, constituting together with the family Prorhynchidae the order Lecithoepitheliata. The pharynx in Gnosonesima represents a type of its own, here called coniform (pharynx coniformis). It has no affinity with the pharynx of the prorhynchids. It is a double structure consisting of an internal plicate pharynx and an external muscular structure comparable with the conorhynch of the Eukalyptorhynchia. G. brattstroemi has epidermal pigment ocelli of a primitive type. Unicellular sensory pits are evidently a common feature in the genus Gnosonesima. The tetrapartite (“tetramerous”) ovary and the genito-intestinal pore are discussed from phylogenetic and taxonomic viewpoints. The order Lecithoepitheliata is evidently polyphyletic but can be retained provisionally for practical reasons.

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