Abstract

Study of Gesneriaceae for the Flora of Ecuador and the Catalogue of the Vascular Plants of Ecuador has shown that some recently published new species need to be transferred with new combinations in Columnea: C. flifera, C. katzensteinii, C. mastersonii, C. medicinalis, C. ornata, and C. tandapiana, or included as synonyms of previously published species. In early 1993 we published a revision of the genus Columnea for Ecuador (Kvist & Skog, 1993). While the 1993 paper was delayed in press by the unfortunate occurrence of Hurricane Iniki in Hawaii, another paper appeared (Wiehler, 1992) that dealt in part with some of the same species that we were describing as new, with names for some of these species under generic names that we consider to be synonyms of Columnea. Therefore, we are here making new combinations for some of the species published in 1992 just prior to our publication. In 1995 additional new species of Columnea and other genera were published by Wiehler, along with further discussion and citation of additional specimens for some of the species published as new in his 1992 paper. We are here making new transfers into Columnea for some of the species occurring in Ecuador and nearby countries published by Wiehler in 1992 and 1995. In most cases we have not seen the holotypes or isotypes because so far as we can determine the specimens have not yet been distributed from GES. However, some of the specimens that we cite as types or representative material in our 1993 paper were cited by Wiehler in his 1992 and 1995 papers. In addition, some of the specimens Wiehler cited in his papers we had also seen. For some of the other species of which we have not seen the types, we are confident from our reading of the descriptions that the species described belong to Columnea or other genera and are here making the transfers or new combinations. Still other species from these papers require future study before we can determine their status. Some of the new species published by Wiehler in 1992 had been transferred to Columnea in the interim before the 1995 papers appeared. However, no mention is made by Wiehler (1995a, 1995b) of the new species or combinations published in 1993 by Smith and Skog or in 1994 by Smith. Columnea filifera (Wiehler) L. E. Skog & L. P. Kvist, comb. nov. Basionym: Dalbergaria filifera Wiehler, Phytologia 73: 224. 1992; Gesneriana 1: 39. 1995. TYPE: Ecuador. Esmeraldas: near new road from Lita and Alto Tambo towards San Lorenzo, Wiehler & GRF Expedition 9033 (holotype, GES not seen; isotypes, QCA, US none seen). Columnea fililoba L. P. Kvist & L. E. Skog, Allertonia 6: 356. 1993. TYPE: Ecuador. Carchi: San Marcos, Kvist, Barfod & Nissen 48997 (holotype, AAU; isotypes, QCA, QCNE, US). Wiehler (1995a: 39) included our 1993 species as a synonym under his Dalbergaria filifera. We agree that the species are synonymous and are therefore making a new combination in Columnea. Columnea katzensteinii (Wiehler) L. E. Skog & L. P. Kvist, comb. nov. Basionym: Pentadenia katzensteinii Wiehler, Phytologia 73: 235. 1992; Gesneriana 1: 75. 1995. TYPE: Ecuador. Morona-Santiago: Cordillera del Boliche, ca. 60 km from Lim6n, Wiehler & GRF Expedition 88128 (holotype, GES not seen; isotypes, F, K, MO, NY, QCA, US none seen). Columnea lavandulacea L. P. Kvist & L. E. Skog, Allertonia 6: 387. 1993. TYPE: Ecuador. Pichincha: Quito-Santo Domingo de los Colorados road, Kvist & Barfod 49066 (holotype, AAU). Duplicates of a specimen cited by Wiehler (1995a: 75) were seen and cited by us as paratypes NovoN 7: 413-416. 1997. This content downloaded from 157.55.39.163 on Wed, 21 Sep 2016 05:11:37 UTC All use subject to http://about.jstor.org/terms

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