Nomenclatural types and taxonomy of Gothan’s Arctic fossil conifer wood

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A century ago Gothan described fossil woods from the Arctic and established several new species and genera that were subsequently widely used. The type material has, however, never been reviewed. Investigations in Berlin and Stockholm resulted in the re-location of the original material which allows us to propose lectotypifications for the names of six species recognized by Gothan, three of which represent types of Gothan's generic names, Anomaloxylon, Protocedroxylon, and Protopiceoxylon. The types (holotypes or lectotypes) are illustrated here, some of them for the first time with micrographs. Taxonomic investigation of Gothan's species with Abietineentupfelungen, an anatomical feature of ray cells, suggests that Cedroxylon transiens and Anomaloxylon magnoradiatum should be regarded as taxonomic synonyms of Protocedroxylon araucarioides, and Cedroxylon cedroides as a taxonomic synonym of Protopiceoxylon exstinctum.