Abstract

Thoracican cirripedes from the Cretaceous (Albian– Maastrichtian) of Europe (UK, France, Germany) and North Africa (Morocco, Tunisia) are described taxonomicallyand significantly aug - ment the diversity previously known from these regions. The new taxa include: a new zeugmatole - padid, Subsecolepas gen. nov. (type species: S. holtwilsoni sp. nov.), three new cretiscalpellid genera, Witherscalpellum gen. nov., Jagtscalpellum gen. nov. and Striascalpellum gen. nov., including the new species J. africanum, J. spinosum, S. bromleyi , S. barringtonensis and S. elegans and a new scalpellid of the genus Regioscalpellum , R. kennedyi. In addition, the Virgiscalpellinae subfam. nov. includes two new genera, Collinslepas and Virgilepas, with the new species C. aitlaminensis, C. robustus , C. tunisiensis, V. aboudaensis, V. peakei and V. hancocki. Paedomorphic evolution in the sub-family led to forms with reduced numbers of capitular plates. Five new species of Virgiscalpellum are also erected: V. mhrilensis sp. nov., V. multilineatum sp. nov., V. truncatum sp. nov., V. laevis sp. nov. and V. sussexiense sp. nov., as well as a new Proverruca, P. anglica. The Eoverrucidae fam. nov. is erected for the genus Eoverruca, for which two new species, E. aubensis (middle Albian, France) and E. symmetrica (upper Campanian, UK) are described. The record from the Albian takes the origin of the verrucomorph cirripedes back 25 million years. Furthermore, two new species of Brachylepas, B. thieli and B. hantonensis, extend the range of the genus down into the Cenomanian, and a new genus of brachylepadid, Fallaxlepas, is erected, with the type species Pollicipes fallax Darwin , 1851. The stratigraphical distribution of Cretaceous cirripede genera is tabulated and demonstrates a remarkable increase in diversity through the Albian to mid-Campanian, above which there is a minor decrease. The Cretaceous– Paleogene (K/Pg) boundary saw the extinction of 15 cirripede genera.

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