New species of Kamopanorpa Martynov from the Permian of South Siberia with comments on the systematic position of Microptysmatidae (Protomeropina = Permotrichoptera)

Abstract

The systematic position of Microptysmatidae and Protomeropidae is critically reviewed in light of the latest reassessment by Minet et al. The Microptysmatidae belongs to a stem-group of Trichoptera, or at least to stem-Amphiesmenoptera (depending on chosen methodology), rather than to its own order Permotrichoptera as suggested by Minet et al., and Protomeropidae are not Mecoptera. The double anal loop of Kamopanorpa Martynov, 1928 is not significantly different from that of more advanced amphiesmenopterans, while in Microptysmella Kukalová-Peck & Willmann, 1990, anal veins show clear tendency to looping (not a “fairly different arrangement”). If Microptysmatidae are considered in the traditional way, the polarity of characters listed by Minet et al. as supporting monophyly of the order Permotrichoptera can be treated just the opposite: divided veins RS1, RS2 and M1 are symplesiomorphies with Protomeropidae, and simple M4 is the synapomorphy of Microptysmatidae, Cladochoristidae, and Amphiesmenoptera s. str. In addition, three new species of Kamopanorpa are described from the Permian deposits of Minusinsk Coal Basin, South Siberia: Kamopanorpa rasnitsyni sp. nov., K. sivchikovi sp. nov., and K. rotunda sp. nov. Some traits of fore- and hindwing venation of Microptysmatidae are discussed.