Abstract
SUMMARY Consideration of the landing patterns of midges (Culicoides spp., Diptera: Ceratopogonidae) on a human subject leads to a generalization of the problem of combining information from a number of samples concerning departure from the Hardy-Weinberg law. Statistics are suggested that generalize those proposed for the original problem. Haldane (1954) and Smith (1970) have proposed closely related statistics for combining the information provided by samples from different populations on deviation from the Hardy-Weinberg law. The problem of testing a hypothesis about insect behaviour leads to a generalization in which the genotypic frequencies for each sample are supplemented by information on gene frequencies. The problem arose from a study of the behaviour of female biting midges (Culicoides spp., Diptera: Ceratopogonidae) carried out near Loch Awe, Argyll, Scotland by Peter Marsh (now deceased) of the Department of Zoology, University of Edinburgh. In order to investigate their temporal landing pattern on a human, he recorded the times at which
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