Middle Permian sponge–microencruster reefal facies in the mid-Panthalassan Akiyoshi atoll carbonates: observations on a limestone slab

Abstract

A Murgabian (Middle Permian) reef-core facies in the Akiyoshi Limestone, southwest Japan, which is of mid-Panthalassan atoll origin, is here described from a limestone slab. The reef-core facies is composed mainly of boundstone with submarine cement and matrix. The reef-building community is characterized by a high-diversity biota, including sponges (sphinctozoans, inozoans, and chaetetids), bryozoans (fistuliporids and cryptostomates), crinoids and various microencrusters (Tubiphytes, Archaeolithoporella, reticular and laminar microbialites, encrusting foraminifers and problematic laminar red algae). Among them, sphinctozoan sponges and microencrusters such as Tubiphytes and microbialites are the most abundant. This community is similar to a time-equivalent lagoonal mound (patch-reef) biota on the Akiyoshi atoll in terms of the dominant sponges, Tubiphytes and microbialites, but differs in the additional occurrence of bryozoans, Archaeolithoporella and encrusting foraminifers, which probably preferred higher-energy conditions in the reef-core environment. Among temporally changing reef-building communities on the Carboniferous–Permian Akiyoshi atolls, the sponge-dominated reefal community described here flourished on mid-Panthalassan atolls in a relatively stable warm-water environment during a Middle Permian post-deglacial period.