Abstract

We explored the effects of heterogeneity of sward height on the functioning of grazing systems through a spatially implicit mechanistic model of grazing and sward growth. The model uses a population dynamic approach where a sward is spatially structured by height, which changes as a function of defoliation, trampling, and growth. The grazing component incorporates mechanisms of bite formation, intake, and digestion rates, but excludes sward quality effects. Sward height selection is determined by maximization of the instantaneous intake rate of forage dry mass. For any given average sward height, intake rate increased with increasing spatial heterogeneity. Spatio-temporal distribution of animal density over paddocks did not markedly affect animal performance but it modified the balance of vegetation heterogeneity within and between paddocks. Herbage allowance was a weak predictor of animal performance because the same value can result from multiples combinations of herbage mass per unit area, number of animals, animal liveweight, and paddock area, which are the proximate determinants of intake rate. Our results differ from models that assume homogeneity and provide strong evidence of how heterogeneity influences the dynamic of grazing systems. Thus, we argue that grazing management and research need to incorporate the concept of heterogeneity into the design of future grazing systems.

Highlights

  • Traditional grazing management has been based on variables such as stocking rate and herbage allowance, aimed at controlling intensity and frequency of defoliation, and herbage production and animal performance

  • Herbage allowance and stocking rate may only describe an instant of a grazing system, but not its dynamics, which result from multiple processes influenced by biotic and abiotic factors

  • We aimed to develop a spatially implicit model of grazing and sward growth to explore the effects of spatial heterogeneity on responses to traditional grazing management variables

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Summary

Introduction

Traditional grazing management has been based on variables such as stocking rate and herbage allowance, aimed at controlling intensity and frequency of defoliation, and herbage production and animal performance. The actual management variables over which we can exert control are shape and area of paddocks, number and characteristics of animals, and timing and duration of grazing and rest periods Additional management variables such as fertilization and irrigation timing and rate can be used to modify the quality and dynamics of vegetation state variables. Livestock distribution and behavior can be altered through rewards like salt or supplements These management and uncontrollable biotic and abiotic factors change over space and time and drive grazing systems into a permanent state of flux

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