Abstract
We develop a fertility model of fitness that is general in that it does not assume that the fitnesses of the mating combinations are symmetrical or that they are additive or multilicative (i. e., that they can be inferred from fitnesses of the two genotypes involved in a mating). %he model considers one locus with three alleles. An experimental test with Drosophila rnelanogaster confirms that the fitnesses of the mating types depart from both additivity (or multiplicativity) and symmetry although this last property is of no consequence for the development of analytical models). urnerical simulations yield the same, or very nearly the same, equilibrium freuencies as the analytical model, independently of whether or not Hardy-Weinberg equilibrium Trequencies are assumed at the beginning of each selection cycle.
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