Abstract

The exploitation of mollusks by the first anatomically modern humans is a central question for archaeologists. This paper focuses on level 8 (dated around ∼ 100 ka BP) of El Harhoura 2 Cave, located along the coastline in the Rabat-Témara region (Morocco). The large quantity of Patella sp. shells found in this level highlights questions regarding their origin and preservation. This study presents an estimation of the preservation status of these shells. We focus here on the diagenetic evolution of both the microstructural patterns and organic components of crossed-foliated shell layers, in order to assess the viability of further investigations based on shell layer minor elements, isotopic or biochemical compositions. The results show that the shells seem to be well conserved, with microstructural patterns preserved down to sub-micrometric scales, and that some organic components are still present in situ. But faint taphonomic degradations affecting both mineral and organic components are nonetheless evidenced, such as the disappearance of organic envelopes surrounding crossed-foliated lamellae, combined with a partial recrystallization of the lamellae. Our results provide a solid case-study of the early stages of the diagenetic evolution of crossed-foliated shell layers. Moreover, they highlight the fact that extreme caution must be taken before using fossil shells for palaeoenvironmental or geochronological reconstructions. Without thorough investigation, the alteration patterns illustrated here would easily have gone unnoticed. However, these degradations are liable to bias any proxy based on the elemental, isotopic or biochemical composition of the shells. This study also provides significant data concerning human subsistence behavior: the presence of notches and the good preservation state of limpet shells (no dissolution/recrystallization, no bioerosion and no abrasion/fragmentation aspects) would attest that limpets were gathered alive with tools by Middle Palaeolithic (Aterian) populations in North Africa for consumption.

Highlights

  • Shell accumulations are found in many MP/MSA archaeological coastal sites in Southern Europe and in Africa [30,31,32,33,34,35,36]

  • The fossil Patella specimens come from level 8 of El Harhoura 2 Cave, Témara, Morocco, from the test pit located in the current cave entrance

  • The fossil shells used in this study seem well preserved and do not present any rounded or polished aspect due to water/sand abrasion

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Summary

Introduction

Shell accumulations are found in many MP/MSA archaeological coastal sites in Southern Europe and in Africa [30,31,32,33,34,35,36] These deposits are mainly composed of mollusk species living on intertidal rocky shores and collected by humans. These species are clearly visible, collectable [31, 37] and concentrated in clusters in comparison to sandy shores mollusk assemblages (with a 14 times higher macro-invertebrate biomass) [38] It seems that such mollusks represented a significant contribution to the diet of human populations inhabiting coastal areas [39]. The shells can be used to trace out harvesting seasons [41, 42], human population densities [43], the exploitation of aquatic resources [44], and the emergence of modern human behaviors [45]

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