Abstract

Comparison of the four species groups comprising the tribe Geissoieae (Lamanonia in South America, Geissois sensu stricto from islands in the south-western Pacific, and Pseudoweinmannia and Geissois in Australia) showed differences in the stipules, inflorescences and flowers. Molecular phylogenetic analyses suggested that Geissois sensu lato is paraphyletic, with Australian Geissois being closer to Pseudoweinmannia than to New Caledonian Geissois. The name Karrabina is published to accommodate the Australian species formerly placed in Geissois, namely K. benthamiana Rozefelds & H.C.Hopkins and K. biagiana Rozefelds & H.C.Hopkins. Distinguishing characters for Karrabina are: four lateral stipules per node that vary from free to largely fused between and sometimes across the petioles; inflorescences with a variable number of metamers and racemes, with both median and lateral modules that always lack a median raceme; and pale flowers with five or six calyx lobes and shortly exserted stamens. Geissois s.s. differs in having: two intrapetiolar stipules per node; inflorescences consisting of few metamers and few racemes, with the modules being always lateral and possessing a median raceme; flowers with four red calyx lobes and prominently exserted stamens. The generic limits of Lamanonia and Pseudoweinmannia remain unchanged.

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