Abstract
Abstract Exclusion of nitrogen and light from existing leaves at initiation of an episode of shoot growth decreased shoot and root relative growth rate. The combination of both nitrogen and light exclusion synergistically impacted relative growth rate for shoot (RGRs) and relative growth rate for root (RGRr). The next episode of shoot growth provided sufficient leaf area for carbohydrate assimilation and maintaining shoot and root growth rates when light was excluded from mature leaves. A better understanding of the developmental and biochemical changes of this episodic species provided useful information for timing of fertilizer application and transplanting of Ligustrum and other episodic species.
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