Abstract

Continental communities represented by complex, rapidly evolving invertebrate ecosystems first appeared during the Early Palaeozoic Era and are recorded in surface and subsurface bioturbation. The Mississippian-age Mauch Chunk Formation contains several high-density bioturbated, fine-grained sandstones that are preserved as erosively based fluvial channel fills. Several of these channel sandstone bodies are almost completely homogenized by subsequent bioturbation throughout their entire thickness, up to a maximum depth of 1.6m.The Mauch Chunk ichnofossil record demonstrates that high-density bioturbation occurred at least 30my prior to the Permian and well before its widespread distribution in the Triassic. Hence, the Late Mississippian high-density bioturbated sandstones demonstrate a much earlier high-density exploitation of the continental subsurface ecospace than previously known.

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