Abstract
Heterochrony is an evolutionary term that describes the comparatively common phylogenetic variation between species in the relative timing of developmental events. Heterochronic variation has also been induced by mutation to identify genes that regulate the timing of developmental events (1, 2). Genes that control the temporal dimension of development, heterochronic genes, can be thought of as the temporal analogs of the homeotic genes, which regulate spatial dimensions (e.g., anterior-posterior, dorsal-ventral axes) during development of metazoans. These pathways generate graded or binary levels of regulatory factors that pattern one axis of the developing animal. Heterochronic genes may be the target of mutations that cause heterochronic change in phylogeny. In the
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