Abstract
Gap frequency of sugar beet and wheat canopies was measured using downward looking hemispherical photography. This technique accounts for gap frequency ( P o ) variation with both zenith θ and azimuth φ angles. A range of leaf area indices was obtained by varying the sowing date, the sowing density or the measurement date. Results show that the exponential law ( P o (θ,φ) = e −K(θ,φ),L ) accurately describes variations with leaf area index ( L) for each direction (θ, φ). Row architecture had very little effect on gap frequency azimuthal variations. A dispersion coefficient (λ o) was directly computed with a Markov chain model applied on sugar beet crops using observed gap frequencies, leaf area indices and leaf inclination. The λ o coefficient value was close to 1.2 for zenith angles below 60°, and then decreased for higher zenith angles. Assuming the dispersion coefficient λ o to be independent of the zenith angle θ allowed us to fit both the λ o value and the average leaf inclination when it was not measured. Fitted values of leaf inclination and λ o indicated that the wheat crop was rather crectophile with a regular leaf arrangement ( λ o ) = 1.56 ). The sugar beet canopy was more planophile and very close to the random leaf arrangement ( λ o = 1.04 ). These results contrasted with those found when λ o was assumed to vary with θ and with the observed leaf inclination value. The discrepancy was partly explained by the fact that the fitted values of λ o absorbed most of the experimental errors and violations of the implicit hypothesis. However, this study provides a simple semi-empirical way to describe gap frequencies of crops throughout their growing cycle or for various densities or sowing dates, using Markov chain processes.
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