FOREST COMMUNITIES IN NORTHWESTERN ALBERTA

Abstract

Spruce, tamarack (larch), balsam fir, pine, and poplar communities of the region are described in terms of floristic composition and ecological relationships. The white spruce (Picea glauca) association is regarded as the climax type of the region. Of four phases or faciations presented by the white spruce association, the feather moss faciation appears to be the climax to which the other faciations tend to develop. Two black spruce (Picea mariana) communities are recognized, the black spruce – feather moss association and the black spruce–peat moss association. Of these, the former is characterized by "feather mosses" such as Hylocomium splendens and has developed on relatively level terrain without much peat formation, whereas the latter has a Sphagnum floor and has arisen in definite depressions through acid bog stages with the production of considerable peat. The black spruce – bog moss community is interpreted as subclimax, with natural succession to the black spruce – feather moss association. The tamarack (Larix laricina) community has many features in common with the black spruce – peat moss association but differs markedly, not only in its dominant species, but because of its development from a Drepano-cladus–Carex–Betula bog under persisting wet conditions. Succession to black spruce commonly occurs. Balsam fir (Abies balsamea) is relatively rare in the region and usually grows in mixed stands with white spruce, paper birch, aspen, and balsam poplar. Two divisions of the pine association are recognized, the jack pine (Pinus banksiana) and the lodgepole pine (P. contorta var. latifolia) consociations. For each of these, two phases are described, the pine – feather moss faciation on the more shaded sites and the pine–heath faciation on the more open and drier areas. Knowledge of the ranges of these two pines in northern Alberta and concerning hybrids between the species is extended. The poplar association, classified as aspen (Populus tremuloides) and balsam poplar (P. balsamifera) consociations, is considered in relation to other vegetation, especially prairie grassland and white spruce. Encroachment of aspen poplar upon native grassland is counteracted by various factors, notably burning. Natural succession of poplar and pine to white spruce is impeded chiefly by forest fires. Some attention is given to phytogeographical problems of this transition region.