Abstract
Using direct-aid climbing techniques for sampling trunks and branch systems, we found 74 species of lichens and 32 species of bryophytes growing as epiphytes in a 450-year-old Douglas fir forest in western Oregon. Six epiphyte zones are described: base, moist side of trunk, dry side of trunk, upper trunk, axes of branch systems and branchlets of branch systems. The flora of each zone is compared with that of the rest of the tree and with that found on understory vegetation. We employed direct-aid mountain-climbing techniques to climb and sample living trees in a Douglas fir forest in western Oregon. Previous descriptions of Douglas fir epiphytes (Szczawinski, 1953; Coleman, Muenscher & Charles, 1956; Hoffman & Kazmierski, 1969) were based on samples from the lower trunk or felled trees. We relate epiphyte distribution to differences in bark and exposure of major sub- divisions of the trunk and canopy. This paper is based on sampling designed to provide estimates of epiphyte biomass per hectare of forest to be used in ecosystem modeling undertaken by the Western Coniferous Biome, U.S. International Biological Program. Qualitative results only are presented here; our sampling methods are outlined to illustrate intensity of sampling. METHODS
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