Abstract
This work was funded by the ANR (Agence National de la Recherche, France) as part of the ‘RALI – The Rise of Animal Life’ project (Grant number ANR-11-BS56-0025), and by the NSF of USA (grant numbers EAR 0921245 and EAR 0922054). Acid treatment and slide preparation were conducted by Laurence Debeauvais (CNRS/Universite Lille 1). Paul K. Strother (Boston College) found the first possible xiphosuran coxae in our material. Massimo Morpurgo (Museum of Nature South Tyrol) provided exuviae of Limulus polyphemus. We thank James C. Lamsdell (Yale University), Petra Tonarova (Tallinn University of Technology, Czech Geological Survey), Olle Hints (Tallinn University of Technology), Denis Audo (Universite de Rennes 1), Carolin Haug (LMU Munich), Joachim T. Haug (LMU Munich) and Alexander Blanke (Hull, UK) for discussing our findings, and two anonymous reviewers for their helpful comments. This work is a contribution to IGCP project 653 – ‘The onset of the Great Ordovician Biodiversification Event’ and to the CPER research project CLIMIBIO. The authors thank the French Ministere de l'Enseignement Superieur et de la Recherche, the Hauts-de-France Region and the European Funds for Regional Economic Development for their financial support to this project and the Department of Innovation, Research and University of the Autonomous Province of Bozen/Bolzano for covering the Open Access publication costs.
Highlights
HAL is a multi-disciplinary open access archive for the deposit and dissemination of scientific research documents, whether they are published or not
Together these methods have yielded exceptionally well-preserved crustacean-type setae and a population of distinctive microfossils which we identify as the feeding appendages of a small-bodied arthropod
The attempt to search for organic microfossils in the Winneshiek Shale has yielded numerous objects that we tentatively identify as crustacean filter plates and coxae of a diminutive euchelicerate, which are described
Summary
HAL is a multi-disciplinary open access archive for the deposit and dissemination of scientific research documents, whether they are published or not. The Winneshiek Shale in Iowa, USA, is a Darriwilian (Middle Ordovician) Konservat-Lagerst€atte (site with exceptionally well-preserved fossils; see Seilacher 1970) that is notable not least for its arthropod fossils. It has yielded the currently oldest described eurypterids (Lamsdell et al 2015a), phyllocarids with softtissue preservation (Liu et al 2006; Briggs et al 2016), ostracods and other bivalved arthropods (Briggs et al 2016), and a dekatriatan-like euchelicerate (Lamsdell et al 2015b). This site is one of few Konservat-Lagerst€atten in the Ordovician (Van Roy et al 2015), the only other Konservat-Lagerst€atte known so far from the Middle Ordovician ( Darriwilian) being the Llanfallteg Formation in south-west
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