Abstract
FENN1, and later Hill2, showed by heat measurements the isolated frog sartorius muscle liberated extra energy in proportion to the mechanical work done by the muscle during contraction (‘Fenn effect’). More recently, analysis of the breakdown of high-energy phosphates also revealed that more energy is utilized by a contracting sartorius muscle, which is allowed to shorten against a load (isotonic contraction), than by a muscle which is not allowed to shorten (isometric contraction)3,4.
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