Abstract
Anthers are tetrasporangiate. Anther wall development is of the dicotyledonous type. The glandular tapetum is single-layered and exhibits dual origin. Microspore tetrads are tetrahedral. Pollen grains at the time of anther dehiscence are two-celled and triaperturate. Ovules are unitegmic, tenuinucellar and anatropous. Development of female gametophyte conforms to the Polygonum type. Fertilization is porogamous. Endosperm is celluar with a single-celled chalazal haustorium, while the micropylar haustorium is initially four-celled, but becomes single-celled and quadrinucleate by the breakdown of the walls of the haustorial cells. Embryo development conforms to Onagrad type. Seed coat is thin with a prominent epidermal layer.
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