Abstract

ABSTRACTMost classical models for the movement of organisms assume that all individuals have the same patterns and rates of movement (for example, diffusion with a fixed diffusion coefficient) but there is empirical evidence that movement rates and patterns may vary among different individuals. A simple way to capture variation in dispersal that has been suggested in the ecological literature is to allow individuals to switch between two distinct dispersal modes. We study models for populations whose members can switch between two different nonzero rates of diffusion and whose local population dynamics are subject to density dependence of logistic type. The resulting models are reaction–diffusion systems that can be cooperative at some population densities and competitive at others. We assume that the focal population inhabits a bounded region and study how its overall dynamics depend on the parameters describing switching rates and local population dynamics. (Traveling waves and spread rates have been studied for similar models in the context of biological invasions.) The analytic methods include ideas and results from reaction–diffusion theory, semi-dynamical systems, and bifurcation/continuation theory.

Highlights

  • The movement of organisms plays an important role in determining the spatial distributions and interactions of populations, and influences many ecological processes

  • Most classical models for the movement of organisms assume that all individuals in a given population have the same patterns and rates of movement but there is empirical evidence that movement rates and patterns may vary among different individuals, or for the same individual under different conditions; see [5, 6, 13, 19, 35, 36, 40]

  • There has been some modelling of populations that can switch between different dispersal modes

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Summary

Introduction

The movement of organisms plays an important role in determining the spatial distributions and interactions of populations, and influences many ecological processes. The results of Girardin [15] show that for a class of models including system (1) but allowing arbitrarily many possible movement modes, if the zero equilibrium is unstable there exist travelling waves connecting the zero equilibrium to some positive equilibrium. (The case of Robin conditions is similar.) Let (λd, φd) be the principal eigenvalue and associated positive eigenfunction of u = λu, x ∈ , u =.

Minimal patch size under Dirichlet boundary conditions
Dynamics under Neumann boundary conditions
Existence of positive equilibria with one component small
Spatially homogeneous case
Conditions for global stability
Bifurcations
Spatially constant solutions
Nonconstant solutions
General boundary conditions
Discussion
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