Abstract

The extent to which tool-using animals take into account the properties of the tool is little explored. The use of percussors to crack open encapsulated fruit is a complex form of tool use, the choice of an adequate tool being a critical aspect in success. Several properties (e.g., material, resistance, friability, shape and weight) affect the suitability of an object to open a hard-shelled nut, with weight being amongst the most important factors. In general, heavier tools require fewer strikes to crack open a nut.

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