Diverse dinosaur-, pterosaur-, and bird-track assemblages from the Hakou Formation, Lower Cretaceous of Gansu Province, northwest China

Abstract

Diverse and well-preserved assemblages of dinosaur (theropod, sauropod and ornithopod), pterosaur, and bird tracks from the Hekou Formation (Lower Cretaceous) in the Yellow River (Huang He) valley represent the first significant fossil footprint discoveries in Gansu Province, China. However, the sites are large, visually spectacular, and well-exposed thanks to labor-intensive hand excavation. The sites have the potential for development as educational and tourist destinations. These sites have become one of the National Geoparks in China. Dinosaur tracks include at least two theropod morphotypes that range in size from about 5 to >30 cm in length. Wide-gauge sauropod tracks ( Brontopodus) range in size from 25 to 90 cm (pes length) and are the best-preserved examples known from China, with clear claw impressions. One trackway suggests an accelerating/running individual. Parallel ornithopod trackways indicate gregarious behavior. An enigmatic trackway may be a manus-only ornithopod trackway. A pterosaur trackway (cf. Pteraichnus), the first reported from China, consists of 24 consecutive footprints, and is the longest, well-preserved trackway on record. Bird tracks (cf. Aquatilavipes) are also very well preserved. The tracks occur at multiple stratigraphic levels in fluvio-lacustrine sequences of paleosol mudstones and sandstones with mud cracks and wave ripple marks. A minimum ichnodiversity of eight, the highest reported from the Cretaceous of China, is estimated. The saurischian component (theropods and sauropods) compares well with Inner Mongolia ichnofaunas from the Jing Chuan Formation. However, the co-occurrence of ornithopod and sauropod tracks is rare in Asia and globally, and compares with assemblages from South Korea, a similar Cretaceous paleolatitude (ca. 30°).