Abstract
SUMMARY The ovule primordium of Averrhoa is trizonate; both integuments originate dermally and are initially mainly 2-layered. The inner integument becomes 5- to 6-layered; its innermost layer is at first endothelium-like. The outer integument is multiplicative. Most cells of the seed coat are crushed. The crystal-bearing endotesta and the fibrous exotegmen constitute the mechanical layers of the seed. The outer walls of the seed epidermis have wall thickenings of a pectic nature covered with a rather thick cuticle. The seed of Sarcotheca is similar to that of Averrhoa. The ruminations are more pronounced and correspond with differences in the dimensions of the adjacent exotegmic cells. The smaller seed of Dapania is somewhat different from those of Averrhoa and Sarcotheca, but also has an oxalidaceous nature. Averrhoa, Sarcotheca and Dapania have to be retained in the Oxalidaceae.
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