Abstract
The dynamically changing land cover configuration and its impact on biodiversity have aroused interest in the study of deforestation and its consequences. Deforestation is generally considered to be one of the most serious threats to biological diversity. Awareness of how different deforestation patterns influence habitat quality of forest patches is essential for efficient landscape–ecological management. The overall effect of deforestation on the forest patch depends on its size, shape and location. Zipperer (1993) identified the following types of the deforestation pattern: Internal deforestation that starts in the forest patch and progresses outwardly; External deforestation that starts outside and cuts into the forest patch, including indentation, cropping and removal; Fragmentation when the patch is split into smaller parcels. Forest fragmentation is one of the most frequently cited causes of species extinction making it a crucial contemporary conservation issue. The classic view of a habitat fragmentation is the breaking up a large intact area of a single vegetation type into smaller landscape units or simply the disruption of continuity (Fahrig, 2003; Lord & Norton, 1990). This process represents a transition from being whole to being broken into two or more distant pieces. The outcome is landscape composed of fragments (e. g. forest) with something else (the nonforest matrix) between the fragments. Fragmentation of biotopes affects several ecological functions of landscape, first of all the spatial distribution of selected plant and animal species and associations (Bruna & Kress; 2002, Kurosawa & Askins, 2003; Parker et al., 2005). Fragments of original biotopes with reduced area and increased isolation are not capable of securing suitable conditions for life and reproduction of some organisms. Consequences of fragmentation include the species biodiversity decline, functional changes in ecological processes, for instance disruption of trophic chains (Valladares et al., 2006), and genetic changes in organisms (Cunningham & Moritz 1998; Gibbs, 2001). The history of research focused on the fragmentation consequences reaches to the 1960s when the theory of insular biogeography was published (MacArthur & Wilson, 1963, as cited in Faaborg et al., 1993). The basis of this theory is the recognition that the smaller and more dispersed islands, the lower number of species is capable to find suitable conditions for their permanent existence. However, the phenomenon is not limited only to islands in the geographic sense. As Madera and Zimova (2004) report similar problems were also
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