Abstract
In this study, developmental modules of modularity and integration of the fore and hind wings of Melanitis leda were described using the MINT software (Modularity and Integration Tool, ver. 1.5, Marquez, 2008). A total of 195 points were used to trace and outline the margins of the wings as well as the major veins. The test was conducted based on a Wishart/Monte Carlo test on a total of 1,000 replicates. The best fit models are determined with the highest P-value and lowest gamma value. Differences in ranks of the top 3 models were observed in both the fore and hind wings of the butterfly. It was hypothesized to be caused by effects associated with the environment, phenotypic plasticity or the signalling sources for position and morphology of pattern elements which only covers short distances.
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