Abstract
A specimen of the fern fructification Radiitheca Brousmiche et al., up to now only known in the Pilsencoalfield (Czech Republic), has been discovered in the late Westphalian D strata of the Sarre Lorraine coalfield. Relationships between Radiitheca dobranyana Brousmiche et al. and Pecopteris waltonii Corsin are discussed both on paleobotanical and palynological basis. TEM study of the Microreticulatisporites (Knox) Potonie & Kremp macerated from the French and Czech specimens led to the characterization of the unique layer of the spore-wall that is the exospore, This investigation has shown: (1) that the ultrastructural features of the spores from both localities are perfectly similar, (2) the nature of the bodies occurring in each lumen that were called “pustules” in Brousmiche et al. (3) and that the lack of these ones is only due to preservation or maceration bias. A comparison of the exospore ultrastructure with that of main groups of living ferns suggests close affinities between the genus Radiitheca and the Gleicheniaceae.
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