Abstract

In 1970 Janzen proposed that species-specific seed predation, by eliminating establishment of seedlings near adult trees of the same species, should lead to a regular dispersion of trees and to increased species diversity in tropical forests. Seed predation would lead to increased species diversity by setting upper bounds to the densities of individual species: if each extant species reached its predation-determined limit and yet more canopy space was available, individuals of additional species would be able to invade. Here I derive a patch model of Janzen's mechanism that is simple in form yet captures its spatial flavor. The model predicts that this mechanism will tend to stabilize any number of competing species. As a corollary, the model predicts that Janzen's mechanism can lead to increased diversity without leading to regular spacing of individuals.

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