Abstract

Whilst an African origin of modern humans is well established, the timings and routes of their expansions into Eurasia are the subject of heated debate, due to the scarcity of fossils and the lack of suitably old ancient DNA. Here, we use high-resolution palaeoclimate reconstructions to estimate how difficult it would have been for humans in terms of rainfall availability to leave the African continent in the past 300k years. We then combine these results with an anthropologically and ecologically motivated estimate of the minimum level of rainfall required by hunter-gatherers to survive, allowing us to reconstruct when, and along which geographic paths, expansions out of Africa would have been climatically feasible. The estimated timings and routes of potential contact with Eurasia are compatible with archaeological and genetic evidence of human expansions out of Africa, highlighting the key role of palaeoclimate variability for modern human dispersals.

Highlights

  • Whilst an African origin of modern humans is well established, the timings and routes of their expansions into Eurasia are the subject of heated debate, due to the scarcity of fossils and the lack of suitably old ancient DNA

  • Analysis of fossil and genetic evidence provides strong support for an African origin of Homo sapiens[1], yet the timing of human expansions out of Africa has been the focus of recent debate[2]

  • While more work is needed to confirm hints of Middle Pleistocene humans in Eurasia, current evidence strongly suggests that Homo sapiens had repeated periods when they were able to leave Africa, the number, timings, routes, and fates of such early waves is unclear

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Summary

Introduction

Whilst an African origin of modern humans is well established, the timings and routes of their expansions into Eurasia are the subject of heated debate, due to the scarcity of fossils and the lack of suitably old ancient DNA. We use high-resolution palaeoclimate simulations of the last 300k years to estimate the tolerance to low precipitation and aridity that would have been required for humans to successfully exit Africa at a given time.

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