Abstract

The Out-of-Africa model holds that anatomically modern humans (AMH) evolved and dispersed from Africa into Asia, and later Europe. Palaeoanthropological evidence from the Near East assumes great importance, but AMH remains from the region are extremely scarce. ‘Egbert’, a now-lost AMH fossil from the key site of Ksar Akil (Lebanon) and ‘Ethelruda’, a recently re-discovered fragmentary maxilla from the same site, are two rare examples where human fossils are directly linked with early Upper Palaeolithic archaeological assemblages. Here we radiocarbon date the contexts from which Egbert and Ethelruda were recovered, as well as the levels above and below the findspots. In the absence of well-preserved organic materials, we primarily used marine shell beads, often regarded as indicative of behavioural modernity. Bayesian modelling allows for the construction of a chronostratigraphic framework for Ksar Akil, which supports several conclusions. The model-generated age estimates place Egbert between 40.8–39.2 ka cal BP (68.2% prob.) and Ethelruda between 42.4–41.7 ka cal BP (68.2% prob.). This indicates that Egbert is of an age comparable to that of the oldest directly-dated European AMH (Peştera cu Oase). Ethelruda is older, but on current estimates not older than the modern human teeth from Cavallo in Italy. The dating of the so-called “transitional” or Initial Upper Palaeolithic layers of the site may indicate that the passage from the Middle to Upper Palaeolithic at Ksar Akil, and possibly in the wider northern Levant, occurred later than previously estimated, casting some doubts on the assumed singular role of the region as a locus for human dispersals into Europe. Finally, tentative interpretations of the fossil's taxonomy, combined with the chronometric dating of Ethelruda's context, provides evidence that the transitional/IUP industries of Europe and the Levant, or at least some of them, may be the result of early modern human migration(s).

Highlights

  • Significant changes in human behaviour, cognition and innovation become sharply evident in the archaeological record of Eurasia at 45,000 years BP and demarcate the end of the Middle Palaeolithic and the onset of the Upper Palaeolithic period

  • The material cultures associated with the latter include the so-called ‘‘transitional’’ technocomplexes, and the subsequent Early Upper Palaeolithic (EUP) technocomplexes, namely the (Proto- and Early) Aurignacian found throughout the continent

  • Comparison of the Ksar Akil specimens with directly and indirectly dated modern humans In the past three decades, several anatomically modern humans (AMH) remains from across Eurasia and Africa have been directly dated between 50–30 ka BP, many of these determinations are potentially problematic due to insufficient decontamination methods used in the past

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Summary

Introduction

Significant changes in human behaviour, cognition and innovation become sharply evident in the archaeological record of Eurasia at 45,000 years BP and demarcate the end of the Middle Palaeolithic and the onset of the Upper Palaeolithic period. With the exception of about 10 teeth from the IUP and Early Ahmarian levels of Ucagızlı Cave in southern Turkey, most likely belonging to Homo sapiens [40,41], Egbert and Ethelruda are the only other human fossils in the Near East directly linked to EUP and IUP assemblages, respectively.

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