Abstract
The basic taxonomy and classification of reptile species and genera often use pholidotic characters. Despite that each species has a standard pattern, there are always deviant individuals in terms of scale number, shape, size, or color. Turtles are excellent models for the study of developmental instability because anomalies are easily detected in the form of malformations, additions, or reductions in the number of scutes or scales. The normal number of carapacial scutes in turtles is five vertebrals, four pairs of costals, and 12 pairs of marginals, a pattern known as “typical chelonian carapacial scutation”. Any deviation of vertebral, costal, or marginal scute numbers or their pattern represents an anomaly. Zangerl & Johnson (1957) documented scutation anomalies in 118 species of turtles belonging to seven families, with higher levels of carapace anomalies in aquatic species compared to semi-aquatic and terrestrial species.
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