Abstract
The development of the free nuclear endosperm ofCapsella was studied using the electron microscope and histochemical procedures. The 2-nucleate and 4-nucleate endosperm contain many chloroplasts, mitochondria, dictyosomes, microbodies, polysomes, and an abundance of RER. There is a sharp decline in the starch and lipid reserves of the central cell during the first few divisions of the primary endosperm nucleus. Different developmental patterns occur in the micropylar and chalazal endosperms. Wall projections grow into the micropylar endosperm from the embryo sac wall and the suspensor wall. An unusual form of SER is continuous with the plasma membrane investing some of the suspensor wall projections of the heart-shaped embryo. The SER branches, forms clear and electron dense vacuoles and forms loops which encircle altered-appearing areas of cytoplasm. The micropylar endosperm of the heart-shaped embryo also contains many chloroplasts, mitochondria, dictyosomes, microbodies, polysomes, stacks of crystal-containing RER, and large nuclei which have many RER connections.
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