Bifurcations and spatiotemporal patterns in a ratio‐dependent diffusive Holling‐Tanner system with time delay

Abstract

In this paper, we concentrate on the spatiotemporal patterns of a delayed reaction‐diffusion Holling‐Tanner model with Neumann boundary conditions. In particular, the time delay that is incorporated in the negative feedback of the predator density is considered as one of the principal factors to affect the dynamic behavior. Firstly, a global Turing bifurcation theorem for τ = 0 and a local Turing bifurcation theorem for τ > 0 are given. Then, further considering the degenerated situation, we derive the existence of Bogdanov‐Takens bifurcation and Turing‐Hopf bifurcation. The normal form method is used to study the explicit dynamics near the Turing‐Hopf singularity. It is shown that a pair of stable nonconstant steady states (stripe patterns) and a pair of stable spatially inhomogeneous periodic solutions (spot patterns) could be bifurcated from a positive equilibrium. Moreover, the Turing‐Turing‐Hopf–type spatiotemporal patterns, that is, a subharmonic phenomenon with two spatial wave numbers and one temporal frequency, are also found and explained theoretically. Our results imply that the interaction of Turing and Hopf instabilities can be considered as the simplest mechanism for the appearance of complex spatiotemporal dynamics.