Abstract
Summary The floral development of Parnassia palustris and Francoa sonchifolia , both aberrant members of Saxifragaceae s.l., was studied in order to determine their taxonomic positions. The findings support the propossal of setting up two independent families, viz. Parnassiaceae and Francoaceae. The floral development of the two species is described below. Parnassia: Quincuncial sepal development; petal primordia well developed; androecium is haplostemonous; epipetal whorl developed in the form of staminodes. The morphogenesis does not support the assumption of a staminate bundle. The development of the gynoecium is of a type of its own showing a complete reduction of the plicate section. The symplicate section develops giving commissural stigmas as well as paracarpous and secondary syncarpous ovary portions. Francoa: Noticeable plastochrones are to be found between the initiation of the whorls. Sepal and petal developments are not inhibited. The androecium is obdiplostemonous (displacement of the epipetal stamen primordia caused by secondary growth of the flower receptacle). The carpels show an impeltate structure and are connected by a centrally placed cone of receptacle at the basal part of the ovary. The sectioning is of a type characteristic of Francoa having both a plicate and a symplicate section.
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