Abstract
Understory vegetation influences several ecosystem services and functions of European beech (Fagus sylvatica L.) forests. Despite this knowledge on the importance of understory vegetation, it is still difficult to measure its three-dimensional characteristics in a quantitative manner. With the recent advancements in terrestrial laser scanning (TLS), we now have the means to analyze detailed spatial patterns of forests. Here, we present a new measure to quantify understory complexity. We tested the approach for different management types, ranging from traditionally and alternatively managed forests and national parks in Germany to primary forests of Eastern Europe and the Ukraine, as well as on an inventory site with more detailed understory reference data. The understory complexity index (UCI) was derived from point clouds from single scans and tested for its relationship with forest management and conventional inventory data. Our results show that advanced tree regeneration is a strong driver of the UCI. Furthermore, the newly developed index successfully measured understory complexity of differently managed beech stands and was able to distinguish scanning positions located on and away from skid-trails in managed stands. The approach enables a deeper understanding of the complexity of understory structures of forests and their drivers and dependents.
Highlights
The understory, i.e., “all vegetation growing under an overstory” [1], is an important functional and structural component of temperate forests
The dataset of tree regeneration was used to verify the performance of the understory complexity index (UCI) with regard to on average a significant regeneration
The suggested understory complexity index (UCI) has proven to yield plausible results distinguishing a variety of stand situations and allows quantitatively comparing these with one another, as presented here for beech forests
Summary
The understory, i.e., “all vegetation growing under an overstory” [1], is an important functional and structural component of temperate forests. Understory vegetation influences ecosystem functions, such as nutrient cycling [2] or biodiversity of stands [3]. It interacts with animal communities [4], e.g., by serving as a food source or shelter, and may impact the future composition of tree species [5]. Ehbrecht et al [10] observed an increase in the overall three-dimensional stand structural complexity due to the presence of dense understory vegetation, such as thickets with shelterwood trees. We need quantitative information on the different elements of the structure of primary forests to serve as a reference [12,13,14]
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