Abstract
Evidence is presented that the asci of Cochliobolus cymbopogonis are partially bitunicate and that both layers of the ascus wall are functional in ascospore liberation. The inelastic ectotunica splits at its apex to allow the extension of the endotunica. The endotunica is incomplete at its base but forms an apical cap which covers the bent tips of the ascospores as they pass through the neck of the pseudothecium. Outside the pseudothecium the ascospores straighten and uncoil, and the cap of the endotunica is thrust aside. Under moist conditions the ascospores may be extruded as a spore tendril several millimetres in length. Such spores can be dispersed by splash.
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