Abstract
The anatomy of 12 pseudococculinid species, representing the two subfamilies and 10 (of 11 known) genera, is described in detail. Three new genera are erected, Yaquinabyssia, Copulabyssia and Amphiplica. Subfamilial and generic systematics are reconsidered and additionally based on anatomical characters. Characters defining subfamilies and genera (Pseudoeoeculininae: Pseudococculina, Notocrater, Tentaoculus, Mcsopelex, Bandabyssia, Kurilabyssia; Caymanabyssiinae: Caymanabyssia, Colotrachelus, Yuquinabyssia, Copulabyssia, Amphiplica) are the modifications in shell and radula, the number and arrangement of gill‐leaflets, the size of the pedal gland, the degree of eye‐reduction and the modifications of the seminal groove and of the copulatory organ. Separation of the Pseudococculinidae Hiekman, 1983 from the Cocculinidae Dall, 1882 at the superfamilial level is confirmed by great differences in their anatomy. Whereas the Cocculinidae are closely related to the Bathysciadiidae (Cocculinoidea). the Lepctcllidae, Pyropeltidae, Pseudococculinidae,? Bathyphytophilidae, Osteopeltidae, Cocculinellidae, Addisoniidae and Choristellidae form a clearly separated supcrfamily (Lepetelloidea). Both superfamilies compose the Cocculiniformia. a primitive but highly specialized arehaeogastropod suborder.
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