Abstract

ABSTRACT A rhinocerotid mandibular remain from the Upper Pliocene Tsubusagawa Formation (around 3.5 Ma) was described and discussed its taxonomic status and paleobiogeographical implication. Compared with Late Miocene to Early Pleistocene and living rhinocerotid species, the studied specimen resembles the mandible of the Pliocene Eurasian taxa of the subtribe Rhinocerotina, Dihoplus and Pliorhinus in having a straight ventral margin, constant mandibular height and probably reduced lower second incisor. This specimen, however, lacks detailed diagnostic characters of these genus. In addition, the taxonomic validity of these taxa has been argued so far. Therefore, this specimen was identified as Rhinocerotina gen. et sp. indet. Japanese Pliocene rhinocerotid records and the present study suggest that the Rhinocerotina was distributed in the central to southwestern part of proto-Japan during the mid-Pliocene. If the studied specimen is truly a species of Dihoplus or Pliorhinus, its close relative might have migrated from northern China as in the case of the ancestor of Japanese Pliocene proboscidean, Stegodon miensis. Further discovery of Japanese Pliocene rhinocerotid specimens could resolve this issue.

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