Additional Notes on the Scrophulariaceae of China

Abstract

Two species, Pterygiella suffruticosa Hong and Scrophularia lhasaensis Hong, and one subspecies, S. chasmophila W. W. Smith subsp. xizangensis Hong, are described as new from southwestern China. In addition, four new combinations are proposed in the genera Triphysaria and Mazus: T chinensis (Hong) Hong, M. pumilus (Burman f.) van Steenis var. wangii (Li) Chin, Mazus pumilus (Burman f.) van Steenis var. delavayi (Bonati) Chin, and Mazus pumilus (Burman f.) van Steenis var. macrocalyx (Bonati) Chin. The following new taxa and combinations are proposed to make the names available for the forthcoming account of the Scrophulariaceae in volume 18 of the Flora of China. Pterygiella suffruticosa Hong, sp. nov. TYPE: China. SW Sichuan: Muli, Ninglang, Chongtianhe, 2000 m, 27 Sep. 1983, Qinghai-Xizang Expedition 14400 (holotype, PE; isotype, PE). Figure 1. A ceteris speciebus differt habitu fruticoso et caulibus teretibus. Shrubs, ca. 1 m tall. Stems terete, gray, 5-7 mm thick; shoots densely pilose. Leaves opposite, sessile, linear-elliptic, 3-5 x 0.3-1 cm, densely puberulent on both surfaces, base cuneate, margin entire, apex acute. Spikes terminal; lower bracts similar to upper leaves in shape and size, gradually decreasing in size upward. Pedicels up to 3 mm long, pilose. Calyx 1.3-1.5 cm long, puberulent, cleft to middle, slightly 2-lipped; lobes subulatedeltoid. Corolla yellow or pale pink, 1.4-1.6 cm long; upper lip galeate, entire; lower lip 3-lobed, middle lobe acute, lateral ones rounded, lanose in 2 vertical patches. Pistil hairy; ovary ellipsoid; style ca. 8 mm long. Pterygiella Oliver, which is endemic to China, consists of five species, all of which are annuals, though P bartschioides Handel-Mazzetti is also a biennial. This new species, P suffruticosa, is readily distinguished from all by its shrubby habit. Paratypes. CHINA. Yunnan: E of Lijiang, banks of the Yangtze River, Rock 10469 (A); Mengtze, in ravine, 1400 m, Henry 9141 (A). Scrophularia lhasaensis Hong, sp. nov. TYPE: China. Xizang (Tibet): Lhasa, hills W of Lhasa and beyond Trisum, 4500 m, on rocks, 1 Sep. 1942, F Ludlow & G. Sherriff 9053 (holotype, A). Scrophulariae chasmophilae W W Smith affinis, sed a qua foliis majoribus, 1.7-4.5 cm longis, 1-2.7 cm latis, integris, corolla minore, ca. 10.5 mm longa, lobis postici labii angustis, 2 mm latis, non imbricatis. Caespitose perennials, less than 15 cm tall, glandular with multicellular hairs throughout except for corolla, stamens, and pistil. Rhizomes ca. 9 mm thick. Stems several, with 1 or 2 pairs of scaly leaves in lower part, branched in upper part. Leaves opposite; petiole 2-3 mm long or upper leaves sessile; leaf blade ovate to ovate-orbicular, 1.7-4.5 x 1-2.7 cm, base rounded or nearly truncate, margin entire or rarely scarcely serrate, apex obtuse or acute. Cymes 2, terminal, forming an inflorescence of 4 flowers; peduncle ca. 1.2 cm long; bracts linear. Pedicel 2-3 mm long. Calyx ca. 5 mm long, glandular-pilose, cleft nearly to base; lobes linear-oblong, obtuse at apex, ca. 1.8 mm wide. Corolla greenish yellow, ca. 10.5 mm long, urceolate, glabrous on both sides; upper lip ca. 3.8 mm long, ca. 2.3 mm longer than lower lip; lobes ca. 2 mm wide, not overlapping; lower lip ca. 2 mm long. Stamens slightly exceeding corolla tube in height, glabrous; staminode clavate. Pistil including the style ca. 8.5 mm long, glabrous, ovary gradually attenuate at apex into style. Scrophularia lhasaensis is most closely related to S. chasmophila W. W. Smith and S. przewalskii Batal. From these, it differs in having larger leaves 1.7-4.5 cm long and 1-2.7 cm wide, entire or subentire leaf margins, smaller corollas ca. 10.5 mm long, and narrower (ca. 2 mm), nonoverlapping lobes of upper corolla lips. NovoN 6: 372-374. 1996. This content downloaded from 207.46.13.129 on Mon, 27 Jun 2016 05:34:17 UTC All use subject to http://about.jstor.org/terms Volume 6, Number 4 Hong 373 1996 Scrophulariaceae of China