Abstract
The Asian freshwater clam, Corbicula fluminea, was emersed at 23°C under three conditions: humid air, N₂ hypoxia, or air with valves clamped or bound shut. Under all conditions, clams became acidotic during emersion. In air, hemolymph pH declined from 790 to 7.14 in 72 h. Hypoxic clams displayed a similar decline in pH, but clams with valves clamped became acidotic more rapidly. Hemolymph CO₂ content (Cco₂) and Pco₂ rose with duration of aerial exposure in all treatment groups. However, hemolymph of bound clams had significantly higher Cco₂ and Pco₂ than that of either air- or N₂-exposed bivalves after 48 h emersion. Although hemolymph Po₂ declined with emersion time in all groups, there was no effect of exposure condition, suggesting that ventilatory behaviors did not affect hemolymph Po₂ or that the effects were masked by rapid utilization of inspired O₂. Hemolymph calcium and osmolality increased with emersion time and were greatest in clams with clamped valves. Presumably, shell calcium carbonate was ...
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