A REVISION OF THE GENUS HOPLANDRIA KRAATZ OF AMERICA NORTH OF MEXICO (COLEOPTERA: STAPHYLINIDAE, ALEOCHARINAE)

Abstract

AbstractA revision of the genus Hoplandria of America north of Mexico is presented. Four subgenera are recognized, of which two (Genosema Notman and Lophomucter Notman) previously were given generic status and one (Arrhenandria) is here newly described. Twelve species are defined, of which eight are described as new: H. alternans, H. isabellae, H. kisatchie, H. klimaszewskii, H. oconee, H. okaloosa, H. sanbornei, and H. smetanai. The species are arranged according to their relationships into species groups within the subgenus Lophomucter, where seven species are assigned. For the remaining subgenera species grouping was not possible due to the smaller number of species. The following synonymies are confirmed (the first specific name is valid): Subg. Hoplandria, H. lateralis (Melsheimer) = H. ochracea Kraatz = H. texana Casey = H. arizonica Casey = H. carinata Casey; Subg. Arrhenandria, H. laeviventris Casey = H. acudentata Dury. The species Genosema sexualis Notman is here synonymized with H. pulchra Kraatz. Hoplandria brittoni Casey was previously regarded as a synonym of H. lateralis (Melsheimer); it is now considered as a junior synonym of Platandria carolinae Casey. Lectotypes are designated for Gyrophaena lateralis Melsheimer, Hoplandria arizonica Casey, H. carinata Casey, H. laeviventris Casey, H. ochracea Kraatz, H. pulchra Kraatz, and H. texana Casey. Keys to subgenera and species are provided. All species are described, diagnostic characteristics of each are illustrated with line drawings and scanning electron microscope photomicrographs, and all available bionomic and distributional data are given. The morphology, systematics, and phylogeny of the subgenera and certain species are discussed, and a checklist of Nearctic species is added.