Abstract
Leaf-size and leaf-shape were followed on expanding leaves of mature sporophytes of four Dryopteris species cultivated for more than 7 years in Sapporo. Blade length(BL, cm), leaf length(LL)and blade width(W)were measured. as an indicator of leaf-size, and BL/W and BL/LL ratios were calculated to generate leaf-shape indices during leaf growth from April to June at 2-8 day intervals. Following BL/W changes as a function of LL increase could be used to characterize different stages of leaf growth between Dryopteris laeta and D. austriaca, and between D. crassirhizoma and D. coreanamontana; however both of these two species had almost similar BL/W and LL on-expanded leaves, respectively. The speed of leaf elongation(dLL/dT, cm/day)was also included for characterization of leaf expansion. Based on relative changes(%)between the maximum and minimum values of BL/W, BL/LL and dLL/dT, six categories could be recognized during the process of leaf expansion.
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