A new species of the eurypterid Hibbertopterus from the Carboniferous of New Mexico, and a review of the Hibbertopteridae

Abstract

ABSTRACT Hibbertopterus lamsdelli sp. nov., from the Late Carboniferous Kinney Quarry Lagerstätte of New Mexico (USA), is a large (ca. 1.1 m long) stylonurid eurypterid (sea-scorpion; Chelicerata), similar to H. scouleri from Scotland but with less serrate segment margins, a wider pretelson, shorter telson (tail-spine), and more parallel ventral keels. It is only the fourth, yet most reliable record of an American hibbertopterid. A taxonomic reassessment of Hibbertopterus regards Dunsopterus and Vernonopterus (but not Cyrtoctenus) as synonyms. Hibbertopterids were aquatic (benthic) scavengers and microphagous sweep-feeders, but their trackways indicate that they were capable of brief terrestrial, seasonal nuptial walks, despite their large size; Hibbertopterus had walking legs with spinose extensions at the base (Laden) to spread their load, and the ventral keels on their telson functioned like sled rails to reduce body drag. Hibbertopterids were interpreted as moving into freshwater during the Late Palaeozoic, but a trackway from the middle Permian Collingham Formation (Ecca Group) of South Africa may be from a marine setting, though further analysis is needed to fully evaluate this possibility.