Abstract
Over recent years, intense field work in caves of Portugal has provided new data on the distribution of subterranean Iberian leiodid beetles. Speonemadus algarvensis sp. nov. is described from caves of southern Portugal. The new species is included in the Speonemadus Jeannel, 1922 escalerai-group (Cholevinae; Anemadini). All species of the S. escalerai-group are revised and S. breuili (Jeannel, 1922) is resurrected as a valid species. A key to identify the species of the S. escalerai-group is provided and the distinctive characters are illustrated. The distribution of all species of the group is mapped with new data together with biogeographic considerations.
Highlights
We describe a new southwestern Iberian species of Speonemadus, that colonizes caves of southern Portugal, and revise all species of the escalerai-group
The life-style of this group is mostly subterranean, they are frequently found under stones or in leaf litter in forests and in caves, walking in bat guano or in the subterranean substrate
Speciation and biogeography of the S. escalerai-group The genus Speonemadus is distributed in the Western Mediterranean area
Summary
Recent intense fieldwork in Portuguese caves has provided abundant material of troglobiont beetles (Reboleira 2007; Reboleira et al 2009, 2010a, 2011b; Reboleira & Ortuño 2011). Among this material, a new species of Speonemadus belonging to the escalerai-group was found. A new species of Speonemadus belonging to the escalerai-group was found This species group comprises three Iberian species: Speonemadus angusticollis (Kraatz, 1870), S. bolivari (Jeannel, 1922) and S. escalerai (Uhagón, 1898). We resurrect Speonemadus breuili (Jeannel, 1922) as a valid species
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