Abstract
Many biomedical systems possess features of multi-level or hierarchical organization. To investigate how structure and behaviour are related in such systems, a simple two level time dependent model is proposed and studied using the methods of non-equilibrium statistical mechanics. Analytic expressions are derived which characterize the model in terms of the intra-level and interlevel couplings, environl1lental conditions, and time scales of interest. The model is shown to possess a continuum of global behaviours, ranging from those of a control hierarchy, wherein one level slaves another, to a hierarchy whcrein each structural level is associated with an essentially autonomous behaviour.
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