Recent decades have witnessed an increased development of schemes for payment for watershed ecosystem services (PWES). However, the public is usually excluded from PWES systems. Reliable and empirical research on PWES from the public perspective is scarce. Aiming to understand public perceptions, attitudes, participation, and responses to PWES, this paper investigated local residents living in the Yongding River watershed area through a face-to-face questionnaire survey. The results showed that the public had limited knowledge of PWES. The public was keen to be involved in PWES decision-making, but the current level of public participation was very low. Regarding willingness to pay (WTP) and willingness to accept (WTA), nearly 55% of the respondents supported paying the upstream residents for protecting the environment if they were beneficiaries in the downstream areas, while 85% of the respondents agreed to accept compensation if they were contributors to environmental improvement in the upstream areas. Although some of the respondents’ daily lives were affected by the watershed environment, they were reluctant to pay, reflecting a sign of “free-riding”. The regression analysis showed that public concerns, values, knowledge of PWES and the watershed environment, and demographic factors determined the WTP and WTA. The results of the contingent valuation method and opportunity costs method showed that the annual payment for headwater conservation areas (Huailai and Yanqing) ranged from CNY 245 to 718 million (USD 36 to 106 million). This study contributes to our limited knowledge and understanding of public sentiment and makes recommendations for improving public receptivity to PWES.
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